HANYUDA's Publications in academic journals and books in English



Akita S., Koiwai K., Hanyuda T., Kato S., Nozaki R., Uchino T., Sakamoto T., Kondo H., Hirono I. and Fujita D. 2018. Development of 11 Ecklonia radicosa (Phaeophyceae, Laminariales) SSRs markers using next-generation sequencing and intra-genus amplification analysis. J. Appl. Phycol. (DOI: 10.1007/s10811-018-1406-5) (in press)

Hanyuda T., Hansen G.I. and Kawai H. 2018. Genetic identification of macroalgal species on Japanese tsunami marine debris and genetic comparisons with their wild populations. Mar. Poll. Bull. (DOI: 10.1016/j.marpolbul.2017.06.053) (in press)

Hanyuda T. and Kawai H. 2018. Genetic examination of the type specimen of Ulva australis suggests that it was introduced to Australia. Phycol. Res. (in press)

Ohtsuka S, Shimono T., Hanyuda T., Shang X., Huang C., Soh H.Y., Kimmerer W., Kawai H., Itoh H., Ishimaru T. and Tomikawa K. 2018. Possible origins of planktonic copepods, Pseudodiaptomus marinus (Crustacea: Copepoda: Calanoida), introduced from East Asia to the San Francisco Estuary based on a molecular analysis. Aquatic Invasions. (in press)


Boedeker C., O'Kelly C., West J.A., Hanyuda T., Neale A., Wakana I., Wilcox M., Karsten U. and Zuccarello G.C. 2017. Molecular phylogeny and taxonomic revision of the genus Wittrockiella (Pithophoraceae, Cladophorales), including the descriptions of W. australis sp. nov. and W. zosterae sp. nov. J. Phycol. 53: 522-540. (DOI: 10.1111/jpy.12530)

Kawai H., Hanyuda T., Gao X., Terauchi M., Miyata M., Lindstrom S.C., Klochkova N.G. and Miller K.A. 2017. Taxonomic revision of the Agaraceae with a description of Neoagarum gen. nov. and reinstatement of Thalassiophyllum. J. Phycol. 53: 261–270. (DOI: 10.1111/jpy.12511)

Ni-Ni-Win, Hanyuda T., Kato A. and Kawai H. 2017. Padina ogasawaraensis and Padina reniformis, two new species of Padina (Dictyotales, Phaeophyceae) from the southern Japan based on morphology and molecular markers. Phycologia 57: 20–31. (DOI: 10.2216/17-25.1)

Terauchi M., Yamagishi T., Hanyuda T. and Kawai H. 2017. Genome-wide computational analysis of the secretome of brown algae (Phaeophyceae). Marine Genomics 32: 49–59. (DOI: 10.1016/j.margen.2016.12.002)


Kawai H., Hanyuda T., Kim S.H., Ichikawa Y., Uwai S. and Peters A.F. 2016. Cladosiphon takenoensis sp. nov. (Ectocarpales s.l., Phaeophyceae) from Japan. Phycol. Res. 64: 212–218. (Doi: 10.1111/pre.12140)

West J.A., Hansen G.I., Hanyuda T. and Zuccarello G.C. 2016. Flora of drift plastics: a new red algal genus, Tsunamia transpacifica (Stylonematophyceae) from Japanese tsunami debris in the northeast Pacific Ocean. Algae 31: 289-301.

Kurihara A., Horiguchi H., Hanyuda T. and Kawai H. 2016. Phylogenetic analysis of mtDNA markers for Asparagopsis taxiformis (Bonemaisoniales, Rhodophyta) provides novel insights into their phylogeography. Phycol. Res. 64: 95-101.

Kawai H., Miyoshi K. and Hanyuda T. 2016. Taxonomic revision of Papenfussiella species (Chordariaceae, Phaeophyceae) in the Northern Hemisphere. Phycologia 55: 308-317.

Kawai T., Hanyuda T., Bolton J. and Anderson R. 2016. Molecular phylogeny of Zeacarpa (Ralfsiales, Phaeophyceae) proposing a new family Zeacarpaceae and its transfer to Nemodermatales. J. Phycol. 52: 682-686.

Kawai H., Kogishi K., Hanyuda T., Arai S., Gurgel C.F., Nelson W., Meinesz A., Tsiamis K. and Peters A.F. 2016. Phylogeographic analysis of the brown alga Cutleria multifida (Tilopteridales, Phaeophyceae) suggests a complicated introduction history. Phycol. Res. 64: 3-10.

Hanyuda T., Heesch S., Nelson W., Sutherland J., Arai S., Boo S.M. and Kawai H. 2016. Genetic diversity and biogeography of native and introduced populations of Ulva pertusa (Ulvales, Chlorophyta). Phycol. Res. 64: 102-109.


Kawai H., Hanyuda T., Draisma S.G.A., Wilce R.T. and Andersen R.A. 2015. Molecular phylogeny of two unusual brown algae, Phaeostrophion irregular and Platysiphon verticillatus, proposal of the Stschapoviales ord. nov. and Platysiphonaceae fam. nov., and a re-examination of divergence times for brown algal orders. J. Phycol. 51: 918-928.

Kawai H., Hanyuda T., Yamagishi T., Kai A., Lane C., McDevit D., Küpper F.C. and Saunders G.W. 2015. Reproductive morphology and DNA sequences of the brown alga Platysiphon verticillatus support the new combination P. gracilis. J. Phycol. 51: 910-917.

Pongparadon S., Zuccarello G.C., Phang S-M., Kawai H., Hanyuda T. and Prathep A. 2015. Diversity of Halimeda (Chlorophyta) from the Thai–Malay Peninsula. Phycologia 54: 349-366.

Kojima R., Hanyuda T. and Kawai H. 2015. Taxonomic re-examination of Japanese Halimeda species using genetic markers, and proposal of a new species Halimeda ryukyuensis (Bryopsidales, Chlorophyta). Phycol. Res. 63: 178-188.

Kawai H., Hanyuda T., Mumford T. and Waaland J.R. 2015. An introduced population of Chorda asiatica (Chordaceae, Laminariales) in Puget Sound, Pacific coast of North America. Phycol. Res. 63: 154-158.


Yang E.C., Peters A.F., Kawai H., Stern R., Hanyuda T., Bárbara T., Müller D.G., Srittmatter M. Prud’Homme van Reine and W.F., Küpper, F.C. 2014. Ligulate Desmarestia (Desmarestiales, Phaeophyceae) revised: D. japonica sp. nov. and D. dudresnayi differ from D. ligulata. J. Phycol. 50: 149-166.

Ni-Ni-Win, Sun Z.-M., Hanyuda T., Kurihara A., Gurgel C.F.D. and Kawai H. 2014. Four newly recorded species of the calcified marine brown macroalgal genus Padina (Dictyotales, Phaeophyceae) for Australia. Aust. Syst. Bot. 26: 448-465.


Kawai H., Hanyuda T., Ridgway L.M. and Holser K. 2013. Ancestral reproductive structure in basal kelp Aureophycus aleuticus. Sci. Rep. 3: 2491.


Kawai H., Kogishi K., Hanyuda T. and Kitayama T. 2012. Taxonomic revision of the genus Cutleria proposing a new genus Mutimo to accommodate M. cylindrica (Cutleriaceae, Phaeophyceae). Phycol. Res. 60: 241–248.

Sun Z.-M., Hanyuda T., Lim P.-E., Tanaka J., Gurgel C.F.D. and Kawai H. 2012. Taxonomic revision of the genus Lobophora (Dicytotales, Phaeophyceae) based on morphological analyses and rbcL and cox3 gene sequences. Phycologia 51: 500-512.

Ni-Ni-Win, Hanyuda T. Draisma S.G.A., Verheij E., Prud'homme van Reine W.F., Lim P.E., Phang S.M. and Kawai H. 2012. Morphological and molecular evidence for two new species of Padina (Dictyotales, Phaeophyceae), P. sulcata and P. calcarea, from the central Indo-Pacific region. Phycologia 51: 576-585.

Okamura H., Yagi M., Kawachi M., Hanyuda T., Kawai H. and Walker I. 2012. Application of rotating cylinder method for ecotoxicological evaluation of antifouling paints. Toxicological and Environmental Chemistry 94: 545-556.


Ni-Ni-Win, Hanyuda T., Draisma S.G.A., Furnari G., Meinesz A., Kawai H. 2011. Padina ditristromatica sp. nov. and Padina pavonicoides sp. nov. (Dictyotales, Phaeophyceae), two new species from the Mediterranean Sea based on morphological and molecular markers. Europ. J. Phycol. 46: 327-341.

Ni-Ni-Win, Hanyuda T., Arai S., Uchimura M. Pranthep A., Draisma S.G.A., Phang S.M., Abbott I., Millar A.J.K. and Kawai H. 2011. A taxonomic study of the genus Padina (Dictyotales, Phaeophyceae) including the descriptions of four new species from Japan, Hawaii, and the Andaman Sea. J. Phycol. 47: 1193-1209.


Peters A.F., van Wijk S.J., Cho G.Y., Scornet D., Hanyuda T., Kawai H., Schroeder D.C., Cock J.M. and Boo S.M. 2010. Reinstatement of Ectocarpus crouaniorum Thuret in Le Jolis as a third common species of Ectocarpus (Ectocarpales, Phaeophyceae) in western Europe, and its phenology at Roscoff, Brittany. Phycol. Res. 58: 157-170.

Ni Ni Win, Hanyuda T., Arai S., Uchimura M., Prethep A., Draisma S.G.A., Soe-Htun U. and Kawai H. 2010. Four new species of Padina (Dictyotales, Phaeophyceae) from the Western Pacific Ocean, and reinstatement of Padina japonica. Phycologia 49: 136-153.


Ni Ni Win, Hanyuda T., Arai S., Uchimura M., Abbott I.A. and Kawai H. 2008. New records of Padina species from the western coast of the Pacific Ocean. Phycol. Res. 56: 288-300.

Kawai H., Hanyuda T., Lindeberg M. and Lindstrom S.C. 2008. Morphology and molecular phylogeny of Aureophycus aleuticus gen. et sp. nov. (Laminariales, Phaeophyceae) from the Aleutian Islands. J. Phycol. 44: 1013-1021.


Kawai H., Shimada S., Hanyuda T., Suzuki T., Gamagori City Office. 2007. Species diversity and seasonal changes of dominant Ulva species (Ulvales, Ulvophyceae) in Mikawa Bay, Japan, deduced from ITS2 rDNA region sequences. Algae 22: 222-230.

Ohtani M., Ohmi T., Uwai S., Hanyuda T., Prabowo R.E., Yamaguchi T. and Kawai H. 2007. Occurrence and diversity of barnacles on international ships visiting Osaka Bay, Japan, and their risk of introduction. Biofouling 23: 277-286.

Lim P.-E., Sakaguchi M., Hanyuda T., Kogame K., Phang S.-M. and Kawai H. 2007. Molecular phylogeny of crustose brown algae (Ralfsiales, Phaeophyceae) iferred from rbcL sequences resulting in the proposal Neoralfsiaceae fam. Nov. Phycologia 46: 456-466.

Kawai H., Hanyuda T., Draisma S.G.A. and Müller D.G. 2007. Molecular phylogeny of Discosporangium mesarthrocarpum (Phaeophyceae) with a reassessment of the order Discosporangiales. J. Phycol. 43: 186-194.


Iima M., Hanyuda T., Ueda K., Yoshizaki M. and Ebata H. 2004. Morphology, ontogeny and phylogeny of freshwater green alga, Blidingia minima var. ramifera (Ulvales, Ulvophyceae) from Nikko, Tochigi Prefecture, Japan. Jap. J. Phycol. 52: 177-182.

Yoshii Y., Hanyuda T., Wakana I., Miyaji, K., Arai S., Ueda K. and Inouye I. 2004. Carotenoid compositions of Cladophora balls (Aegagropila linnaei) and some members of the Cladophorales (Ulvophyceae, Chlorophyta): their taxonomic and evolutionary implication. J. Phycol. 40: 1170-1177.

Hanyuda T., Suzawa Y., Arai S., Ueda K. and Kumano S. 2004. Phylogeny and taxonomy of freshwater Bangia (Bangiales, Rhodophyta) in Japan. J. Jap. Bot. 79: 262-268.

Hanyuda T., Suzawa Y., Suzawa T., Arai S., Sato H., Ueda K. and Kumano S. 2004. Biogeography and taxonomy of Batrachospermum helminthosum Bory (Batrachospermales, Rhodophyta) in Japan inferred from rbcL gene sequences. J. Phycol. 40: 581-588.


Hanyuda T., Wakana I., Arai S., Miyaji K., Watano Y. and Ueda K. 2002. Phylogenetic relationships within Cladophorales (Ulvophyceae, Chlorophyta) inferred from 18S rRNA gene sequences, with special reference to Aegagropila linnaei. J. Phycol. 38: 564-571.


Hanyuda T., Arai S. and Ueda K. 2000. Variability in the rbcL Introns of Caulerpalean Algae (Chlorophyta, Ulvophyceae). J. Plant Res. 113: 403-413.


Ueda K., Hanyuda T., Nakano A., Shiuchi T., Seo A., Okubo H. and Hotta M. 1997. Molecular phylogenetic position of Podostemaceae, A marvelous aquatic flowering plant family. J. Plant Res. 110: 87-92.


Kawai H., Hanyuda T. and Uwai S. 2016. Evolution and biogeography of laminarialean kelps. (Ed. Hu Z.M. and Fraser C.) pp. 227-249. In 'Seaweed Phylogeography - Adaptation and Evolution of Seaweeds under Envirionmental Change’, Springer.

Ni-Ni-Win, Hanyuda T. Draisma S.G.A., Lim P.E., Phang S.M. and Kawai H. 2013. Taxonomy of the genus Padina (Dictyotales, Phaeophyceae) based on morphological and molecular evidences, with key to species identification. In Taxonomy of Southeast Asian Seaweeds II (Phang S.M. and Lim P.E. (Eds.)). Institute of Ocean and Earth Sciences, University of Malaya Monograph Series 15. University of Malaya Press, Kuala Lumpur.