KAWAI's Publications in academic journals and books in English

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Suzuki, M., Terada, R., Shibata, K., Kawai, H. 2021. New records of Chondracanthus saundersii and Schottera koreana (Gigartinales, Rhodophyta) from Japan based on molecular and morphological analyses. Phycol. Res. Published online. 10.1111/pre.12447

Vieira, C., Steen, F., D’hondt, S., Bafort, Q., Tyberghein, L., Fernandez- García, C., Wysor, B., Tronholm, A., Mattio, L., Payri, C., Kawai, H., Saunders, G.W., Leliaert, F., Verbruggen, H., De Clerck, O. 2021. Global diversification and biogeography of a group of brown seaweeds (Phaeophyceae) driven by clade-specific evolutionary processes. J. Biogeography Published online. 10.1111/jbi.14047

Ni-Ni-Win, Mya-Kyawt-Wai, Geraldino, P. J. L., Liao, L. M., Aye, C.-T. P. P., Ni Ni Mar, Hanyuda, T., Kawai, H., Tokeshi, M. 2021. Taxonomy and species diversity of the brown algal genus Padina (Dictyotales, Phaeophyceae) from the Indo-Pacific with the description of two new species. Europ. J. Phycol. Published online. 10.1080/09670262.2021.1883742

Vieira, C., Akita, S., Uwai, S., Hanyuda, T., Shimada, S., Kawai, H. 2021. Hildenbrandia (Hildenbrandiales, Florideophyceae) from Japan and taxonomic lumping of H. jigongshanensis and H. japananensis. Phycol. Res. Published online 10.1111/pre.12456

Akita, S., Koiwai, K., Ishikawa, T., Sakamoto, T., Yoshimura, T., Kyomoto, S., Nanri, K., Kato, Y., Krashima, A., Hanyuda, T., Shimada, S., Kawai, H., Fujita, D. 2021. Molecular evidence for naturally occurring intra and intergeneric hybridization in Ecklonia (Laminariales, Phaeophyceae). Phycologia, published online. 10.1080/00318884.2021.1885246

Vieira, C., N’Yeurt, A.D.R., Faravavy, R.A., D’Hondt, S., Tran, L.A. Van den Spiegel, D., Kawai, H., De Clerck, O. 2021. Marine macroalgal biodiversity of northern Madagascar: morpho-genetic systematics and implications of anthropic impacts for conservation. Biodiversity & Conservation, 30: 1501–1546.

Vieira, C., Faravavy, R.A., Zubia, M., Bolton, J.J., Anderson, R.J., Engelen, A.H., D’Hondt, S., Leliaert, F., Payri, C., Kawai, H., De Clerke, O. 2021. Lobophora (Dictyotales) from the western Indian Ocean: diversity and biogeography. South African J. Botany. In press.

Kawai, H., Watanabe, Y., Kawai, H. 2021. Taxonomic revision of Saundersella (Ectocarpales s.l., Phaeophyceae) with description of Saundersella crassa sp. nov. and transfer of Heterosaundersella hattoriana to Saundersella. Phycologia published online. 10.1080/00318884.2021.1916859

Vieira, C., Faravavyc, R. A. Zubiad, M. Engelene, A. H., D’Hondtb, S., Kawai, H., De Clerck, O. 2021. Lobophora (Dictyotales, Phaeophyceae) from the western Indian Ocean: diversity and biogeography. South Afr. J. Bot. In press.


Hanyuda, T., Yamamura, K., Kawai, H. 2020. Molecular studies of Gloiopeltis (Endocladiaceae, Gigartinales), with recognition of G. compressus comb. nov. from Japan. Phycologia 59: 1–5. 10.1080/00318884.2019.1663476

Hanyuda, T., Yamamura, K., Boo, G.H., Miller, K.A., Vinogradova, K.L., Kawai, H. 2020. Identification of true Gloiopeltis furcata (Gigartinales, Rhodophyta) and preliminary analysis of its biogeography. Phycol. Res.68: 161–168. 10.1111/pre.12411

Kawai, H., Akita, S., Hashimoto, K. and Hanyuda, T. 2020. A multigene molecular phylogeny of Eisenia reveals evidence for a new species, Eisenia nipponica (Laminariales), from Japan. Europ. J. Phycol. 55: 234–241. 10.1080/09670262.2019.1692911

Kawai, H., Terada, R., Hanyuda, T. 2020. Molecular phylogeny of Punctaria mageshimensis reveals evidence for its transfer to Spatoglossum as S. mageshimensis (Dictyotales, Phaeophyceae). Phycol. Res. 68: 203–207. 10.1111/pre.12417

Kawai, H., Hanyuda, T., Akita, S., Uwai, S. 2020. The macroalgal culture collection in Kobe University (KU-MACC), and a comprehensive molecular phylogeny of macroalgae based on the culture strains. Appl. Phycol. Published online (open access). 10.1080/26388081.2020.1745685

Rizouli, A., Küpper F. C., Louizidou, P., Mogg, A. O. M., Azzopardi, E., Sayer, M. D. J., Kawai, H., Hanyuda, T., Peters A.F. 2020. The minute chromophyte alga Schizocladia ischiensis (Schizocladiophyceae, Ochrophyta) raised by germling emergence from substratum collected at 24 m depth off Rhodes (Dodecanese, Greece). Diversity 2020, 12(3), 102. 10.3390/d12030102

Kawai, H., Hanyuda, T., Mine, I., Takaichi, S., Terada, R., Kitayama, T. 2020. Morphology and molecular phylogeny of Umbraulva spp. (Ulvales, Ulvophyceae), and proposal of Ryuguphycus gen. nov. and R. kuaweuweu comb. nov. Europ. J. Phycol. Published online 10.1080/09670262.2020.1753815

Akita, S., Hashimoto, K., Hanyuda, T., Kawai, H. 2020. Molecular phylogeny and biogeography of Ecklonia spp. (Laminariales, Phaeophyceae) in Japan revealed taxonomic revision of E. kurome and E. stolonifera. Phycologia 59: 333–339. 10.1080/00318884.2020.1756123

Bringloe, T., Starko, S. Wade, R.M., Vieira, C., Kawai, H., De Clerck, O., Cock, M., Coelho, S., Destombe, C., Valero, M., Neiva, J., Pearson, G.A., Faugeron, S., Serrão, E.A., Verbruggen, H. 2020. Phylogeny and evolution of the brown macroalgae. Critical Reviews in Plant Sciences. Published online. 10.1080/07352689.2020.1787679

NiNiWin, Hanyuda, T., Kato, A., Shimabukuro, H., Uchimura, M., Kawai, H., Tokeshi, M. 2020. Global species diversity and geographical distribution of the genus Padina (Dictyotales, Phaeophyceae): new insights based on molecular and morphological analyses. J. Phycol. in press


Kawai, H., Hanyuda, T., Shibata, K., Kamiya, M., Peters, A.F. 2019. Proposal of a new brown algal species Mesogloia japonica sp. nov. (Chordariaceae, Phaeophyceae) and transfer of Sauvageaugloia ikomae to Mesogloia. Phycologia 58: 63–69. 10.1111/pre.12367

Hanyuda, T., Takeuchi, K., Kawai, H. 2019. Tinocladia sanrikuensis sp. nov. (Ectocarpales s.l., Phaeophyceae) from Japan. Phycol. Res. 67: 221–227. 10.1111/pre.12367

Kawai, H., Hanyuda, T., Sun, Z.M, Bárbara, I., Peters, A.F. 2019. Taxonomic revision of Eudesme (Ectocarpales s.l., Phaeophyceae) proposing a new species E. borealis sp. nov. Phycologia 58: 351–358. 10.1080/00318884.2019.1578630

Kawai, H., Suzuki, M., Saunders, G.W. and Hanyuda, T. 2019. Taxonomic study of the brown algal genus Chorda (Chordaceae, Laminariales) with description of the new species Chorda borealis from Alaska and northern Canada. Europ. J. Phycol. 54: 509–517. 10.1080/09670262.2019.1610580

Kinoshita-Terauchi, N., Shiba, K., Terauchi, M., Romero, F.,Ramírez-Gómez, H., Yoshida, M., Motomura, T., Kawai, H., Nishigaki, T. 2019. High potassium seawater inhibits ascidian sperm chemotaxis, but does not affect the male gamete chemotaxis of a brown alga. Zygote

Starko, S., Martone, P., Keeling, P., Kawai, H., Soto, M., Soto, Lindstrom, M., Darby H., Graham, S., Yotsukura, N., Demes, K. 2019. A comprehensive kelp phylogeny sheds light on the evolution of an ecosystem. Mol. Phylog. Evol. 136: 138–150. 10.1016/j.ympev.2019.04.012

Terada, R., Abe, M., Abe, T., Aoki, M. Dazai, A., Endo, H., Kamiya, M., Kawai, H., Kurashima, A., Motomura, T., Murase, N., Sakanishi, Y., Shimabukuro, H., Tanaka, J., Yoshida, G., Aoki, M. 2019. Japan’s nationwide long-term monitoring survey of seaweed communities known as the “Monitoring Sites 1000”: Ten-year overview and future perspectives. Phycol. Res. published online 10.1111/pre.12395

Hanyuda, T., Aoki, S., Kawai, H. 2019. Reinstatement of Myelophycus caespitosus (Scytosiphonaceae, Phaeophyceae) from Japan. Phycol. Res. 68:126–134. 10.1111/pre.12405

Zanolla, M., Carmosa, R., Kawai, H., Stengel, G., Altamirano, M. Role of thermal photosynthetic plasticity in the dispersal and settlement of two global green tide formers: Ulva pertusa and U. ohnoi. Mar. Biol. 166:123. 10.1007/s00227-019-3578-1


Hanyuda, T., Kawai, H., 2018. Genetic examination of the type specimen of Ulva australis suggests that it was introduced to Australia. doi: 10.1111/pre.12222[PDF]

Yamaguchi, A., Wakeman, K.C., Hoppenrath, M., Horiguchi, T., Kawai, H. 2018. Molecular phylogeny of the benthic dinoflagellate Cabra matta (Dinophyceae) from Okinawa, Japan. Phycologia. 57: 630–640. 10.2216/18-7.1

Hansen, G.I., Hanyuda, T., Kawai, H. 2018. The invasion threat of benthic marine algae arriving on Japanese tsunami marine debris (JTMD) in Oregon and Washignton, USA. Phycologia. 57: 641–658. 10.2216/18-58.1

NiNiWin, Hanyuda, T., Kato, A., Kawai, H. 2018. Two new species of Padina (Dictyotales, Phaeophyceae) from southern Japan, P. ogasawaraensis sp. nov. and P. reniformis sp. nov., based on morphology and molecular markers. Phycologia 57: 20–31. DOI:10.2216/17-25.1

Ohtsuka, S., Shimono, T., Hanyuda, T., Shang, H., Huang, C., Soh, X.Y., Kimmerer, M., Kawai, H., Itoh, H., Ishimaru, T., Tomikawa, K. 2018. Possible origins of planktonic copepods, Pseudodiaptomus marinus (Crustacea: Copepoda: Calanoida), introduced from East Asia to the San Francisco Estuary based on a molecular analysis. Aquatic Invasions 13: 221–230. https://doi.org/10.3391/ai.2018.13.2.04


Kawai, H., Hanyuda, T., Gao, X., Terauchi, M., Miyata, M., Lindstrom, S.C., Klochkova, N.G., Miller, K.A. 2017. Taxonomic revision of the Agaraceae with a description of Neoagarum gen. nov. and reinstatement of Thalassiophyllum. J. Phycol. 53: 261-70. DOI: 10.1111/jpy.12511[PDF]

Calvyn F.A. Sondak, C.F.A., Ang, P.O.Jr., Beadall, J., Bellgrove, A., Boo, S.M., Gerung, G.S., Hepburn, C.D., Hong, D.D., Hu, Z., Kawai, H., Lim, P.-E., Largo, D., Lee, J.A., Mayakun, J., Nelson, W.A., Oak, J.H., Phang, S.M., Sahoo, D., Peerapornpis, Y., Yang, Y., Chung, I.K. 2017. Carbon dioxide mitigation potential of seaweed aquaculture beds (SABs). J. Appl. Phycol. DOI: 10.1007/s10811-016-1022-1[PDF]

Terauchi, M., Yamagishi, T., Hanyuda, T., Kawai, H. 2017. Genome-wide computational analysis of the secretome of brown algae (Phaeophyceae). Marine Genomics. 32: 49–59. doi.org/10.1016/j.margen.2016.12.002[PDF]

Poong, S.-W, Lim P.-E., Phang, S.-M., Sunarpi, H., West, J.A., Miller, K.A., Nelson, W.A., Kawai, H. 2017. Two new species of Mesospora (Ralfsiales, Phaeophyceae) from the subtropical Indo-Pacific region. Phycologia 56: 487–498. DOI: 10.2216/16-142.1.

Hanyuda, T., Hansen, G.I., Kawai, H. 2017. Genetic identification of macroalgal species on Japanese tsunami marine debris and genetic comparisons with their wild populations. Mar. Poll. Bull. 10.1016/j.marpolbul.2017.06.053[PDF]


Yamaguchi, A., Yoshimatsu, S., Hoppenrath, M., Wakeman, K.C., Kawai, H. 2016. Molecular phylogeny of the benthic dinoflagellate genus Amphidiniopsis and its relationships with the family Protoperidiniaceae. Protist 167: 568-583. doi.org/10.1016/j.protis.2016.09.003[PDF]

Suzuki, M., Segawa, T., Mori, H., Akiyoshi, A., Ootsuki, R., Kurihara, A., Sakayama, H., Kitayama, T., Abe, T., Kogame, K., Kawai, H., Nozaki, H. 2016. Next-generation sequencing of an 88-year-old specimen of the poorly known species Liagora japonica (Nemaliales, Rhodophyta) supports the recognition of Otohimella gen. nov. PLOS ONE DOI: 10.1371/journal.pone.0158944.[PDF]

Kawai, H., Hanyuda, T., Kim, S.H., Ichikawa, Y., Uwai, S., Peters, A.F. 2016. Cladosiphon takenoensis sp. nov. (Ectocarpales s.l., Phaeophyceae) from Japan. Phycol. Res. in press[PDF]

Kawai, H., Hanyuda, T., Bolton, J., Anderson, R. 2016. Molecular phylogeny of Zeacarpa (Ralfsiales, Phaeophyceae) proposing a new family Zeacarpaceae and its transfer to Nemodermatales. J. Phycol. 52:682-686[PDF]

Hanyuda, T., Heesch, S., Nelson, W., Sutherland, J., Arai, S., Boo, S.M., Kawai, H. 2016. Genetic diversity and biogeography of native and introduced populations of Ulva pertusa (Ulvales, Chlorophyta). Phycol. Res. 64: 102–109.[PDF]

Kurihara, A., Horiguchi, H., Hanyuda, T., Kawai, H. 2016. Phylogenetic analysis of mtDNA markers for Asparagopsis taxiformis (Bonemaisoniales, Rhodophyta) provides novel insights into their phylogeography. Phycol. Res. 64: 95–101.[PDF]

Kawai, H., Miyoshi, K., Hanyuda, T. 2016. Taxonomic revision of Papenfussiella species (Chordariaceae, Phaeophyceae) in the Northern Hemisphere. Phycologia 55: 308–317. DOI: 10.2216/15-139.1[PDF]

Kawai, H., Kogishi, K., Hanyuda, T., Arai, S., Gurgel, C.F., Nelson, W., Meinesz, A., Tsiamis, K., Peters, A.F. 2016. Phylogeographic analysis of the brown alga Cutleria multifida (Tilopteridales, Phaeophyceae) suggests a complicated introduction history. Phycol. Res. 64: 3–10. DOI: 10.1111/pre.12113.[PDF]


Fu, G., Nagasao, C., Yamagishi, T., Kawai, H., Okuda, K., Horiguchi, T., Motomura, T. 2015. Ubiquitous distribution of helmchrome in phototactic swarmers of the stramenopiles. Protoplasma, DOI 10.1007/s00709-015-0857-7[PDF]

Yamagishi, T., Kurihara, A., Kawai, H. 2015. A ribbon-like structure in the ejective organelle of the green microalga Pyramimonas parkeae (Prasinophyceae) consists of core histones and polymers containing N-acetyl-glucosamine. Protist 166: 522-533. doi:10.1016/j.protis.2015.08.003[PDF]

Kawai, H., Hanyuda, T., Draisma, S.G., Wilce, R.T., Andersen, R.A. 2015 Molecular phylogeny of two unusual brown algae, Phaeostrophion irregulare and Platysiphon glacialis, proposal of the Stschapoviales ord. nov. and Platysiphonaceae fam. nov., and a re-examination of divergence times for brown algal orders. J. Phycol. 51: 918-928. DOI: 10.1111/jpy.12332

Kawai, H., Hanyuda, T., Yamagishi, T., Kai, A., Lane, C.E., McDevit, D., Küpper, F.C., Saunders, G.W. 2015. Reproductive morphology and DNA sequences of the brown alga Platysiphon verticillatus support the new combination Platysiphon glacialis J. Phycol. 51: 910-917. DOI: 10.1111/jpy.12331

Pongparadon, S., Zuccarello, G.C., Phang, S.-M., Kawai, H., Hanyuda, T., Prathep, A. 2015. Diversity of Halimeda (Chlorophyta) from the Thai-Malay Peninsula. Phycologia. 54: 349-366.

Kojima, R., Hanyuda, T., Kawai, H. 2015. Taxonomic re-examination of Japanese Halimeda species using genetic markers, and proposal of a new species Halimeda ryukyuensis (Bryopsidales, Chlorophyta). Phycol. Res. 63: 178–188.

Sakayama, H., Kai, A., Nishiyama, M., Watanabe, M.M., Kato, S., Ito, M., Nozaki, H., Kawai, H. 2015. Taxonomy, morphology and genetic variation of Nitella flexilis var. bifurcata (Charales, Characeae) from Japan. Phycol. Res. 63: 159–166.[PDF]

Kawai, H., Hanyuda, T., Mumford, T., Waaland, J.R. 2015. An introduced population of Chorda asiatica (Chordaceae, Laminariales) in Puget Sound, Pacific coast of North America. Phycol. Res. 63: 154–158.


Toyoshima, M., Yamagishi, T., Aikawa, S., Kondo, A., Kawai, H. 2014. A pilot-scale floating closed culture system for the multicellular cyanobacterium Arthrospira platensis NIES-03. J. Appl. Phycol. 27: 2191-2202. DOI 10.1007/s10811-014-0484-2[PDF]

Kato, S., Kawai, H., Takimoto, M., Suga, H., Yohda, K., Horiya, K., Higuchi, S., Sakayama, H. 2014. Occurrence of an endangered species Nitellopsis obtusa (Charales, Charophyceae) in western Japan and the genetic differences within and among Japanese populations. Phycol. Res. 62: 222–227.[PDF]

Yamagishi, T., Müller, D.G., Kawai, H. 2014. Comparative transcriptome analysis of Discosporangium mesarthrocarpum (Phaeophyceae), Schizocladia ischiensis (Schizocladiophyceae) and Phaeothamnion confervicola (Phaeothamniophyceae), with special reference to cell wall-related genes. J. Phycol. 50: 543-551. DOI: 10.111/jpy.12190

Kawai, H. 2014. Recent advances in the phylogeny and taxonomy of Laminariales. Perspectives in Phycology. ISSN 2198-011X (online) 1: 27-40.

Ni-Ni-Win, Zhong-Min Sun, Z.-M., Hanyuda, T., Kurihara, A., Gurgel, C.F.D., Kawai, H. 2014. Four newly recorded species of the calcified marine brown macroalgal genus Padina (Dictyotales, Phaeophyceae) for Australia. Aust. Syst. Bot. 26: 448-465. DOI: 10.1071/SB13025[PDF]

Kawai, H., Kitamura, A., Mimura, M., Mimura, T., Tahara, T., Aida, D., Sato, K., H. Sasaki. 2014. Radioactive cesium accumulation in seaweeds by the Fukushima 1 Nuclear Power Plant accident – two years’ monitoring at Iwaki and its vicinity. J. Plant Res. 127: 23-42. doi:10.1007/s10265-013-0603-1. [PDF]

Yang, E.C., Peters, A.F., Kawai, H., Stern, R., Hanyuda, T., Bárbara, T., Müller, D.G., Srittmatter, M. Prud’Homme van Reine, W.F., Küpper, F.C. 2014. Ligulate Desmarestia (Desmarestiales, Phaeophyceae) revised: D. japonica sp. nov. and D. dudresnayi differ from D. ligulata. J. Phycol. 50: 149-166. DOI: 10.1111/jpy.12148


Kawai, H., Hanyuda, T., Ridgway, L.M., Holser, K. 2013. Ancestral reproductive structure in basal kelp Aureophycus aleuticus. Sci. Rep. 3, 2491; DOI:10.1038/srep02491. ISSN (online): 2045-2322.

Aikawa, S., Joseph, A., Yamada, R., Izumi, Y., Yamagishi, T., Matsuda, F., Kawai, H., Chang, J-S., Hasunuma, T., Kondo, A. 2013. Direct conversion of Spirulina to ethanol without pretreatment of enzymatic hydrolysis processes. Energy Environ. Sci. 6: 1844–1849. DOI: 10.1039/C3EE40305J

Poong, S.-W., Lim P.-E., Phang, S.-M., Sunarpi, H., West, J.A., Kawai, H. 2013. A molecular-assisted floristic survey of crustose brown algae (Phaeophyceae) from Malaysia and Lombok Island, Indonesia based on rbcL and partial cox1 genes. J. Appl. Phycol. 26: 1231–1242. DOI 10.1007/s10811-013-0081-9. Online: 20 July 2013.[PDF]

Poong, S.-W., Lim P.-E., Phang, S.-M., Gerung G., Kawai, H. 2013. Mesospora elongata sp. nov. (Ralfsiales, Phaeophyceae), a new crustose brown algal species from the Indo-Pacific region. Phycologia 52: 74–81. DOI: 10.2216/12-42.1[PDF]


Heesch, S., Day, J.G., Yagagishi, T., Kawai, H., Müller, D.G., Küpper, F.C. 2012. Cryopreservation of the model alga Ectocarpus (Phaeophyceae). CryoLetters 33(5): 327–336.[PDF]

Kawai, H., Kogishi, K., Hanyuda, T., Kitayama, T. 2012. Taxonomic revision of the genus Cutleria proposing a new genus Mutimo to accommodate M. cylindrica (Cutleriaceae, Phaeophyceae). Phycol. Res. 60: 241–248.

Yamagishi, T., Kawai, H. 2012. Cytoskeleton organization during the cell cycle in two stramenopile microalgae, Ochromonas danica (Chrysophyceae) and Heterosigma akashiwo (Raphidophyceae), with special reference to F-actin organization and its role in cytokinesis. Protist 163: 686–700.

Blunden, G., Guiry, M.D., Druehl, L.D., Kogame, K., Kawai, H. 2012. Trigonelline and other betains in species of Laminariales. Nat. Prod. Comm.7(7): 863-865.

Sun, Z.-M., Hanyuda, T., Lim, P.-E., Tanaka, J., Gurgel, C.F.D., Kawai, H. 2012. Taxonomic revision of the genus Lobophora (Dicytotales, Phaeophyceae) based on morphological analyses and rbcL and cox3 gene sequences. Phycologia 51: 500-512.[PDF]

Yamagishi, T., Kai, T., Kawai, H. 2012. Trichocyst ribbons of a cryptomonads are constituted of homologs of R-body proteins produced by the intracellular parasitic bacterium of Paramecium. Mol. Evol. 74:147-157. doi: 10.1007/s00239-012-9495-2

Ni-Ni-Win, Hanyuda, T. Draisma, S.G.A., Prud'homme van Reine, W.F., Lim, P.E., Phang, S.M., Kawai, H. 2012. Morphological and molecular evidence for two new species of Padina (Dictyotales, Phaeophyceae), P. sulcata and P. calcarea, from the central Indo-Pacific region. Phycologia 51: 576-585.[PDF]

Okamura, H., Yagi, M., Kawachi, M., Hanyuda, T., Kawai, H., Walker, I. 2012. Application of rotating cylinder method for ecotoxicological evaluation of antifouling paints. Toxicol. Env. Chem. 94: 545-556. DOI: 10.1080/02772248.2012.662811


Phillips, N., Kapraun, D., Garreta, A.G., Ribera, Siguan, A.R., Rull, J.L., Soler, N.S., Lewis, R., Kawai, H. 2011. Estimates of nuclear DNA content in ninety eight species of brown algae (Phaeophyta). AoB. Plants doi: 10.1093/aobpla/plr001[PDF]

Ni-Ni-Win, Hanyuda, T., Arai, S., Uchimura, M. Pranthep, A., Draisma, S.G., Phang, S.M., Abbott, I., Millar, A.J.K., Kawai, H. 2011. A taxonomic study of the genus Padina (Dictyotales, Phaeophyceae) from the subtropical Western Pacific and Indo-West Pacific regions with descriptions of four new species. J. Phycol. 47: 1193-1209.[PDF]

Ni-Ni-Win, Hanyuda, T., Draisma S.G.A., Furnari G., Meinesz A., Kawai, H. 2011. Padina ditristromatica sp. nov. and Padina pavonicoides sp. nov. (Dictyotales, Phaeophyceae), two new species from the Mediterranean Sea based on morphological and molecular markers. Europ. J. Phycol. 46: 327-341. DOI: 10.1080/09670262.2011.614355

Yamagishi, T., Kawai, H. 2011. Cortical F-Actin reorganization and a contractile ring-like structure found during the cell cycle in the red cryptomonad, Pyranomonas helgolandii. J. Phycol. 47: 1121-1130.

Kamiya, M., Kawai, H., Moon, D., Goff, L.J. 2011. Isolation and characterization of phase-specific cDNAs from carposporophytes of Gracilariopsis andersonii (Gracilariales, Rhodophyta). Europ. J. Phycol. 46: 27-35. DOI: 10.1080/09670262.2010.548101


Draisma, S.G.A., Prud’homme van Reine, W., Kawai, H. 2010. A revised classification of the Sphacelariales (Phaeophyceae) inferred from a psbC and rbcL based phylogeny. Europ. J. Phycol. 45: 308-326. DOI: 10.1080/09670262.2010.490959

Peters, A.F., van Wijk, S.J., Cho, G.Y., Scornet, D., Hanyuda, T., Kawai, H., Schroeder, D.C., Cock, J.M., Boo, S.M. 2010. Reinstatement of E. crouaniorum Thuret in Le Jolis as a third common species of Ectocarpus (Ectocarpales, Phaeophyceae) in western Europe, and its phenology at Roscoff, Brittany. Phycol. Res. 58: 157-170.

Cock, J.M., Sterck, L., Rouzé, P., Scornet, D., Allen A.E., Amoutzias, G., Anthouard, V., Artiguenave, F., Aury, J.M., Badger, J.H., Beszteri, B., Billiau, K., Bonnet, E., Bothwell, J.H.F., Bowler, C., Boyen, C., Brownlee, C., Carrano, C.J., Charrier, B., Cho, G.Y., Coelho, S.M., Collén, J., Corre, E., Delage, L., Delaroque, N., Dittami, S.M., Doulbeau, S., Elias, M., Farnham, G., Gachon, C.M.M, Gschloessl, B., Heesch, S., Jabbari, K., Jubin, C., Kawai, H., Kimura, K., Kloareg, B., Küpper, F.C., Lang, D., Bail, A.L., Leblanc, C., Lerouge, P., Lohr, M., Lopez, P.J., Martens, C., Maumus, F., Michel, G., Miranda-Saavedra, D., Morales, J., Moreau, H., Motomura, T., Nagasato, C., Napoli, C.A., Nelson, D.R., Nyvall-Collén, P., Peters, A.F., Pommier, C., Potin, P., Poulain, J., Quesneville, H., Read, B., Rensing, S.A., Ritter, A., Rousvoal, S., Samanta, M., Samson, G., Schroeder, D.C., Ségurens, B., Strittmatter, M., Tonon, T., Tregear, J., Valentin, K., von Dassow, P., Yamagishi, T., Van de Peer, Y., Wincker, P. 2010. The Ectocarpus genome and the independent evolution of multicellularity in the brown algae. Nature 465: 617-621.

Kogishi, K., Kitayama, T., Miller, K. A., Hanyuda, T., Kawai, H. 2010. Phylogeography of Cutleria cylindrica (Cutleriales, Phaeophyceae) in northeastern Asia, and the identity of an introduced population in California. J. Phycol. 46: 553-558.

Ni-Ni-Win, Hanyuda, T., Arai, S., Uchimura, M., Prethep, A., Draisma, S.G.A., Soe-Htun, U., Kawai, H. 2010. Four new species of Padina (Dictyotales, Phaeophyceae) from the Western Pacific Ocean, and reinstatement of Padina japonica. Phycologia 49: 136-153.[PDF]

Tanaka A., Uwai, S., Nelson, W. and Kawai, H. 2010. Phaeophysema gen. nov. and Vimineoleathesia gen. nov., new brown algal genera for the minute Japanese members of the genus Leathesia. Europ. J. Phycol. 45: 109-117. DOI: 10.1080/09670260903383271


Yamagishi, T., Motomura, T., Nagasato, C., Kawai, H. 2009. Novel proteins comprising the stramenopile tripartite mastigoneme in Ochromonas danica (Chrysophyceae). J. Phycol. 45: 1110-1115.

Henkel, S. K., Kawai, H., Hofmann, G. 2009. Interspecific and interhabitat variation in hsp70 gene expression in native and invasive kelp populations. Mar. Ecol. Prog. Ser. 386: 1-13.

Yamaguchi, T., Prabowo, R. E., Ohshiro, Y., Shimono, T., Jones, D., Kawai, H., Otani, M., Oshino, A., Inagawa, S., Akaya, T., Tamura, I. 2009. The introduction to Japan of the Titan barnacle, Megabalanus coccopoma (Darwin, 1854) (Cirripedia: Balanomorpha) and the role of shipping in its translocation. Biofouling. 25: 325-333.

Niwa, K., Iida, S., Kato, A., Kawai, H., Kikuchi, N., Kobiyama, A., Aruga, Y. 2009. Genetic diversity and introgression in two cultivated species (Porphyra yezoensis and P. tenera) and closely related wild species of Porphyra (Bangiales, Rhodophyta). J. Phycol. 45: 493-502.

Uwai, S., Kogame, K., Yoshida, G., Kawai, H., Ajisaka, T. 2009. Geographical genetic structure and phylogeography of the Sargassum horneri/filicinum complex in Japan, based on the mitochondrial cox3 haplotype. Marine Biology 156:901-911.

Kato, A., Guimarães, S.M.P.B., Kawai, H., Masuda, M. 2009. Characterization of the crustose red alga Peyssonnelia japonica (Rhodophyta, Gigartinales) and its taxonomic relationship with P. boudouresquei based on morphological and molecular data. Phycol. Res. 57: 74-86.


Ni-Ni-Win, Hanyuda, T., Arai, S., Uchimura, M., Abbott, I.A., Kawai, H. 2008. New records of Padina species from the western coast of the Pacific Ocean. Phycol. Res. 56: 288-300.

Kawai, H., Hanyuda, T., Lindeberg, M., Lindstrom, S.C. 2008. Morphology and molecular phylogeny of Aureophycus aleuticus gen. et sp. nov. (Laminariales, Phaeophyceae) from the Aleutian Islands. J. Phycol. 44: 1013-1021.

Ikuta, K., Kawai, H., Müller, D.G., Ohama, T. 2008. Recurrent invation of mitochondrial group II introns in specimens of Pylaiella littoralis (brown alga), collected worldwide. Current Genetics 53: 207-216.

Tang, J.C., Xiao Y., Oshima, A., Kawai, H., Nagata, S. 2008. Disposal of seaweed wakame (Undaria pinnatifida) in composting process by marine bacteria Halomonas sp. AW4. Int. J. Biotechnology 10: 73-85.

Niwa, K, Kato, A., Kobiyama, A., Kawai, H., Aruga, Y. 2008. Comparative study on wild and cultivated Porphyra yezoensis (Bangiales, Rhodophyta) based on molecular phylogeny and morphology. J. Appl. Phycol. 20: 261-270.


Kawai, H., Shimada, S., Hanyuda, T., Suzuki, T., Gamagori City Office. 2007. Species diversity and seasonal changes of dominant Ulva species (Ulvales, Ulvophyceae) in Mikawa Bay, Japan, deduced from ITS2 rDNA region sequences. Algae 22: 222-230.

Shimono, T., Iwasaki, N., Kawai, H. 2007. A new species of Dactylopusioides (Copepoda: Harpacticoida: Thalestridae) infesting brown algae, and its life history. Zootaxa 1582: 59-68.

Cho, G.Y., Kogame, K., Kawai, H., Boo, S.M. 2007. Genetic diversity of Scytosiphon lomentaria (Scytosiphonaceae, Phaeophyceae) from the Pacific and Europe based on RuBisCO large subunit and spacer, and ITS nrDNA sequences. Phycologia 46: 657-665.[PDF]

Lim, P.E., Sakaguchi, M., Hanyuda, T., Kogame, K., Phang, S.-M., Kawai, H. 2007. Molecular phylogeny of crustose brown algae (Ralfsiales, Phaeophyceae) inferred from rbcL sequences resulting in proposal for Neoralfsiaceae fam.nov. Phycologia 46: 456-466.[PDF]

Aguilar-Rosas, L.E., Aguilar-Rosas, R., Kawai, H., Uwai, S., Valenzuela-Espinoza. E. 2007. New record of Sargassum filicinum Harvey (Fucales, Phaeophyceae) in the Pacific Coast of Mexico. Algae 22: 17-21.

Kawai, H., Kamiya, M., Komatsu, T., Nakaoka, M., Yamamoto, T., Marine Research Group of Takeno. 2007. Ten years’ monitoring of intertidal macroalgal vegetation of Hyogo Prefecture, northwestern coast of Honshu, Japan to assess the impact of the Nakhodka oil spill. Algae 22: 37-44.

Ohtani, M., Ohmi, T., Uwai, S., Hanyuda, T., Prabowo, R.E., Yamaguchi, T., Kawai, H. 2007. Occurrence and diversity of barnacles on international ships visiting Osaka Bay, Japan, and their risk of introduction. Biofouling 23: 277-286.

Uwai, S., Arai, S., Morita, T., Kawai, H. 2007. Genetic distinctness and phylogenetic relationships among Undaria spp. based on mitochondrial cox3 sequence. Phycol. Res. 45: 687-695.

Yamagishi, T., Motomura, T., Nagasato, C., Kato, A., Kawai, H. 2007. A tubular mastigoneme-related protein ocm1 isolated from the flagellum of a chromophyte alga Ochromonas danica. J. Phycol. 43: 519-527.

Tanaka, A., Nagasato, C., Uwai, S., Motomura, T., Kawai, H. 2007. Reexamination of ultrastructures in the stellate chloroplasts of brown algae; Splachnidium rugosum, Scytothamnus australis, Asteronema ferruginea and Asterocladon interjectum (Phaeophyceae). Phycol. Res. 55: 203-213.

Yamada, M., Yamamoto, K., Ushihara, Y., Kawai, H. 2007. Variation in metal concentrations in the brown alga Undaria pinnatifida in Osaka Bay, Japan. Phycol. Res. 55: 222-230.

Tang, J.C., Wei, J.-H., Maeda, K., Kawai, H., Zhou, Q., Hosoi-Tanabe, S., Nagata, S. 2007. Degradation of seaweed wakame (Undaria pinnatifida) by composting process with inoculation of Bacillus sp. HR6. Biocontrol Science 12: 47-54.

Ajisaka, T., Kim, S.-H., Uwai, S., Kawai, H. 2007. Cladosiphon umezakii sp. nov. (Ectocarpales, Phaeophyceae) from Japan. Phycol. Res. 55: 203-213.[PDF]

Miller, K. A., Engle, J., Uwai, S., Kawai, H. 2007. First report of the Asian seaweed Sargassum filicinum Harvey (Fucales) in California, USA. Biological Invasions 9: 609-613.

Kawai, H., Hanyuda, T., Draisma, S.G.A., Müller, D.G. 2007. Molecular phylogeny of Discosporangium mesarthrocarpum (Phaeophyceae) with a reassessment of the order Discosporangiales. J. Phycol. 43: 186-194.

Sasaki, H., Kawai, H. 2007. Taxonomic revision of the genus Chorda (Chordaceae, Laminariales) based on sporophyte anatomy and molecular phylogeny. Phycologia 46: 10-21.[PDF]


Kato, A., Baba, M., Kawai, H., Masuda, M. 2006. Reassessment of the little-known crustose red algal genus Polystrata Heydrich (Gigartinales), based on morphology and SSU rDNA sequences. J. Phycol. 42: 922-933.

Uwai, S., Nelson, W., Neill, K., Wang, W. D., Aguilar-Rosas, L.E., Boo, S.M., Kitayama, T., Kawai, H. 2006. Genetic diversity in Undaria pinnatifida (Laminariales, Phaeophyceae) deduced from mitochondria genes – origins and succession of introduced populations. Phycologia 45: 687-695.[PDF]

Sasaki, H., Ishida, A., Hashimoto, Y., Takeda, S., Oshima, A. Kawai, H., Nagata, S. 2006. Utilization of proline in Escherichia coli K-12 at different osmolarities. J. Biol. Sci. 6: 675-679.

Yamada, M., Yamamoto, K., Ushihara, Y., Kawai, H. 2006. A survey of organic solvent extractable metal concentrations in the bottom sediment in Osaka Bay, Japan. Mar. Poll. Bull. 52: 214-238.

Yano, T., Kamiya, M., Murakami, A., Sasaki, H., Kawai, H. 2006. Biochemical phenotypes corresponding to molecular phylogeny of the red algae Plocamium (Plocamiales, Rhodophyta): implications of incongruence with the conventional taxonomy. J. Phycol. 42: 155-169.


Kato, A., Masuda, M., Kawai, H. 2005. New records of Peyssonnelia armorica and P. harveyana (Rhodophyta, Gigartinales) from Japan. Phycol. Res. 53: 266-274.

Sasaki, H., Murakami, A., Kawai, H. 2005. Inorganic ion compositions in the Ulvophyceae and Rhodophyceae, with special reference to sulfuric acid ion accumulations. Phycol. Res. 53: 134-137.

Sasaki, H., Murakami, A., Kawai, H. 2005. Seasonal stability of sulfuric acid accumulation in the Dictyotales (Phaeophyceae). Phycol. Res. 53: 85-92.

Fujita, S., Iseki, M., Yoshikawa, S., Makino, Y., Watanabe, M., Motomura, T., Kawai, H., Murakami, A. 2005. Identification and characterization of a fluorescent flagellar protein from the brown alga Scytosiphon lomentaria (Scytosiphonales, Phaeophyceae): A flavoprotein homologous to Old Yellow Enzyme. Eur. J. Phycol. 40:159-167.

Kawai, H., Sasaki, H., Maeba, S., Henry, E.C. 2005. Morphology and molecular phylogeny of Phaeostrophion irregulare (Phaeophyceae) with proposal of Phaeostrophiaceae fam. nov., and review of Ishigeaceae. Phycologia 44: 169-182.[PDF]


Posudin, Y., Kamiya, M., Murakami, A., Kawai, H. 2004. Effect of high irradiances on chlorophyll fluorescence of Ulva pertusa. International Journal on Algae 6: 235-250.

Altamirano, M., Murakami, A., Kawai, H. 2004. High light stress in the kelp Ecklonia cava. Aquatic Botany 79: 125-135.

Yoshikawa, S., Nagasato, C., Makino, Y., Murakami, A., Kawai, H., Ichimura, T., Motomura, T. 2004. Nuclear histone proteins of gametes in brown algae. Jpn. J. Phycol. 52 (Suppl.): 123-127.

Kawai, H., Sasaki, H. 2004. Morphology, life history, and molecular phylogeny of Stschapovia flagellaris (Tilopteridales, Phaeophyceae), and the erection of the Stschapoviaceae fam. nov. J. Phycol. 40: 1156-1169.

Shimono, T., Iwasaki, N., Kawai, H. 2004. A new species of Dactylopusioides (Copepoda: Harpacticoida: Thalestridae) infesting a brown alga, Dictyota dichotoma in Japan. Hydrobiologia 523: 9-15.

Yano. M., Kamiya, M., Arai, S., Kawai, H. 2004. Morphological homoplasy in the Japanese Plocamium species (Plocamiales, Rhodophyta) inferred from the Rubisco spacer sequence and intracellular acidity. Phycologia 43: 383-393.

Sasaki, H., Kataoka, H., Murakami, A., Kawai, H. 2004. Sulfuric acid ion accumulations in brown algae, with special reference to sulfuric acid ion accumulations. Hydrobiologia 512: 255-262.


Nagasato, C., Yoshikawa, S., Yamashita, M., Kawai, H., Motomura, T. 2003. Pyrenoid formation associated with the cell cycle in the brown alga, Scytosiphon lomentaria (Scytosiphonales, Phaeophyceae) J. Phycol. 39: 1172-1180.

Sasaki, H., Lindstrom, S.C., Waaland, J.R., Kawai, H. 2003. Occurrence of the gametophyte of Agarum clathratum (Laminariales, Phaeophyceae) as an endophyte in Orculifilum denticulatum (Gigartinales, Rhodophyceae). Phycol. Res. 51: 192-202.

Kawai, H., Maeba, S., Sasaki, H., Okuda, K., Henry, E. 2003. Schizocladia ischiensis: a new filamentous marine chromophyte belonging to a new class, Schizocladiophyceae. Protist 154: 211-228.

Altamirano, M., Murakami, A., Kawai, H. 2003. Photosynthetic performance and pigment contents of different developmental stages of Ecklonia cava (Laminariales, Phaeophyceae). Bot. Mar. 46: 9-16.

Yamamoto, T., Nakaoka, M., Komatsu, T., Kawai, H., Marine Life Research Group of Takeno, Ohwada, K. 2003. Impacts of the heavy-oil spill from the Russian tanker NAKHODKA on intertidal ecosystems: recovery of animal community. Marine Pollution Bulletin 47: 91-98.

Komatsu, T., Nakaoka, M., Kawai, H., Yamamoto, T., Marine Life Research Group of Takeno, Ohwada, K. 2003. Impacts of the NAKHODKA heavy-oil spill on an intertidal ecosystem: an approach to impact evaluation with geographical information system. Marine Pollution Bulletin 47: 99-104.

Kim, Sung-Ho, Peters, A. F., Kawai, H. 2003. Taxonomic revision of Sphaerotrichia divaricata (C. Ag.) Kylin (Ectocarpales s.l., Phaeophyceae), with a reappraisal of S. firma (Gepp) A. Zinova from the Northwestern Pacific. Phycologia 42: 183-192.[PDF]


Murakami, A., Kawai, H., Adachi, K., Sakawa, T., Miyashita, H., Mimuro, M. 2002. A novel chlorophyll d -containing organism: Discovery and its significance. J. Photoscience 9: 74-77.

Yoshikawa, Y., Nagasato, C., Makino, Y., Murakami, A., Kawai, H. Ichimura, T., Motomura, T. 2002. Nuclear histone proteins of gametes in an oogamous and two isogamous brown algae. J. Phycol. 38: 318-324.

Kim, Sung-Ho, Kawai, H. 2002. Taxonomic revision of Chordaria flagelliformis (Chordariales, Phaeophyceae) with elevation of f. chordaeformis to an independent species. Phycologia 41: 328-339.[PDF]

Flores-Moya, A, Posudin, Y. I., Fernandez, J. A., Figueroa, F. L., Kawai, H. 2002. Photomovement of the swarmers of the brown algae, Scytosiphon lomentaria and Petalonia fascia: effect of photon irradiance, spectral composition and UV dose. J. Photochem. Photobiol. B. Biology 66: 134-140.

Kamiya, M., Kawai, H. 2002 Depedence of the carposporophyte on the maternal gametophyte in three ceramiacean algae (Rhodopyta), with respect to carposporophyte development, spore productiom and germination success. Phycologia 41:107-115.[PDF]


Sasaki, H., Flores-Moya, A., Henry, E.C., Müller, D.G., Kawai, H. 2001. Molecular phylogeny of Phyllariaceae, Halosiphonaceae and Tilopteridales (Phaeophyceae). Phycologia 40: 123-134.[PDF]


Kawai, H., Sasaki, H., Maeda, Y., Arai, S. 2000. Morphology, life history and molecular phylogeny of Chorda rigida sp. nov. (Laminariales, Phaeophyceae) from the Sea of Japan and the genetic diversity of Chorda filum. J. Phycol. 37: 130-142.[PDF]

Kawai, H., Sasaki, H. 2000. Molecular phylogeny of the brown algal genera Akkesiphycus and Halosiphon (Laminariales), resulting in the circumscription of the new families Akkesiphycaceae and Halosiphonaceae. Phycologia 39: 416-428.[PDF]

Kamiya, M., West, J. A., Zuccarello, G. C., Kawai, H. 2000. Caloglossa intermedia sp. nov. (Delesseriaceae, Rhodophyta) from the western Atlantic coast: molecular and morphological analyses with special reference to C. leprieurii and C. monosticha. J. Phycol. 39: 411-420.


Kamiya, M., Tanaka, J., King, R.J., West, J.A., Zuccarello, G.C., Kawai, H. 1999. Reproductive and genetic distinction between broad and narrow entities of Caloglossa continua (Delesseriaceae, Rhodophyta). Phycologia. 38: 356-367.[PDF]

Sasaki, H., Kamiya, M., Kataoka, H., Kawai, H. 1999. Accumulation of sulfuric acid in Dictyotales (Phaeophyceae): Taxonomic distribution and ion chromatography of cell extracts. J. Phycol. 35: 732-739.[PDF]

Boo, S.M, Lee, W.J., Yoon, H.S., Kato, A., Kawai, H. 1999. Molecular phylogeny of Laminariales (Phaeophyceae) inferred from small subunit ribosomal DNA sequences. Phycol. Res. 47: 109-114.[PDF]

Horiguchi, T., Kawai, H., Kubota, M., Takahashi, T., Watanabe, M. 1999. Phototactic responses of four marine dinoflagellates with different types of eyespot and chloroplast. Phycol. Res. 47: 101-107.[PDF]

Chi, Eun-Sup, Henry, E.C., Kawai, H., Okuda, K. 1999. Immunogold-labeling analysis of alginate distributions in the cell walls of chromophyte algae. Phycol. Res. 47: 53-60.[PDF]

Kawai, H., Prud'homme van Reine, W.F. 1998. Life history of Japanese Stypocaulon durum (Sphacelariales, Phaeophyceae). Phycol. Res. 46: 263-270.[PDF]

Ehara, M, Watanabe, K.I., Kawai, H., Inagaki, Y., Hayashi-Ishimaru, Y., Ohama, T. 1998. Distribution of the mitochondrial deviant genetic code, AUA for methionine, in heterokont algae. J. Phycol. 34: 1005-1008.[PDF]

Maier, I., Wolf, S., Delaroque, N., Müller, D.G., Kawai, H. 1998. A DNA virus infecting the marine brown alga Pilayella littoralis (Ectocarpales, Phaeophyceae) in culture. Europ. J. Phycol. 33: 213-220.[PDF]

Kawai, H. 1997. Morphology and life history of Coelocladia arctica (Dictyosiphonales, Phaeophyceae), new to Japan. Phycol. Res. 45: 183-187.[PDF]

Maier, I., Rometsch, E., Wolf, S., Kapp, M., Müller D.G., Kawai, H. 1997. Passage of a marine algal DNA virus from Ectocarpus fasciculatus (Ectocarpales, Phaeophyceae) to Myriotrichia clavaeformis (Dictyosiphonales, Phaeophyceae): Infection symptoms and recovery. J. Phycol. 33: 838-844.[PDF]

Kawai, H., Nakayama, T., Inouye, I., Kato, A. 1997. Linkage of 5S ribosomal DNA to other rDNAs in the chromophytic algae and related taxa. J. Phycol. 33: 505-511.[PDF]

Kogame, Y., Kawai, H. 1996. Development of the intercalary meristem in Chorda filum (Laminariales, Phaeophyceae) and other primitive Laminariales. Phycol. Res. 44: 247-260.[PDF]

Yamano, K. Saito, H., Ogasawara, Y. Fujii, S., Yamada, H. Shirahama, H. and Kawai, H. 1996. The autofluorescent substance in the posterior flagellum of swarmers of the brown alga Scytosiphon lomentaria. Zeitschrift für Naturforschung (c) 51: 155-159.[PDF]

Kawai, H., Nakamura, S. Mimuro, M., Furuya M. and Watanabe, M. 1996. Microspectrofluorometry of the autofluorescent flagellum in phototactic brown algal zoids. Protoplasma 191(3/4): 172-177.[PDF]

Kawai, H. and Tokuyama, M. 1995. Laminarionema elsbetiae gen. et sp. nov. (Ectocarpales, Phaeophyceae), a new endophyte in Laminaria sporophytes. Phycol. Res. 43 (4): 185-190.[PDF]

Kawai, H., Muto, H., Fujii, T. and Kato, A. 1995. A linked 5S rRNA gene in Scytosiphon lomentaria (Scytosiphonales, Phaeophyceae). J. Phycol. 31(2): 306-311.


Kawai, H., Boland, W. and Müller, D.G. 1994. Sexual reproduction and sexual pheromones in Myelophycus simplex (Harvey) Papenfuss (Phaeophyceae). Jpn. J. Phycol. 42: 227-231.

Kawai, H. 1993. Morphology and life history of Kurogiella saxatilis gen. et sp. nov. (Chordariales, Phaeophyceae). Phycologia 32: 462-467.[PDF]

Peters, A.F., Kawai, H. and Novaczek, I. 1993. Intraspecific sterility barrier confirms that introduction of Sphaerotrichia divaricata (Phaeophyceae, Chordariales) into the Mediterranean was from Japan. Hydrobiologia 260/261: 31-36.[PDF]

Kogame, K. and Kawai, H. 1993. Morphology and life history of Japanese Petalonia zosteriforia (Reinke) O. Kunze (Scytosiphonales, Phaeophyceae) from Japan. Jpn. J. Phycol. 41: 29-37.

Schmid, C.E., Schroer, N., Kawai, H. and Müller, D.G. 1992. Isolation and biochemical characterization of different gamete membranes in the chromophyte alga Ectocarpus siliculosus (Phaeophyceae). Plant Physiol. Biochem. 30: 703-712.

Kawai, H. 1992. A summary of the morphology of chloroplasts and flagellated cells in the Phaeophyceae. Kor. J. Phycology. 7: 33-43.

Kitayama, T., Kawai, H., and Yoshida, T. 1992. Dominance of female gametophytes in field populations of Cutleria cyrindrica (Cutleriales, Phaeophyceae) in the Tsugaru Strait, Japan. Phycologia 31: 449-461.[PDF]

Kawai, H. 1992. Green flagellar autofluorescence in brown algal swarmers and their phototactic responses. Bot. Mag. Tokyo. 105: 171-184.[PDF]

Kawai, H. 1991. Critical review of the taxonomy and life history of Kjellmania arasakii (Dictyosiphonales, Phaeophyceae). Jpn. J. Phycol. 39: 319-328.

Kitayama, T., Kawai, H. and Yoshida, T. 1991. Morphological observations on Sphacelaria califorica Sauvageau ex Setchell et Gardner (Sphacelariales, Phaeophyceae) new to Japan. Jpn. J. Phycol. 39: 183-187.

Müller, D.G. and Kawai, H. 1991. Sexual reproduction of Ectocarpus siliculosus (Ectocarpales, Phaeophyceae) in Japan. Jpn. J. Phycol. 39: 151-155.

Kreimer, G., Kawai, H., Müller, D.G. and Melkonian, M. 1991. Reflective properties of the stigma in male gametes of Ectocarpus siliculosus (Phaeophyceae) studied by confocal laser scanning microscopy. J. Phycology 27: 268-276.

Kawai, H., Kubota, M., Kondo, T. and Watanabe, M. 1991. Action spectra of the phototaxis in zoospores of a brown alga Pseudochorda gracilis. Protoplasma 161: 17-22.[PDF]

Kawai, H. and Yamada, I. 1990. The specific identity and life history of Japanese Syringoderma (Syringodermatales, Phaeophyceae). Bot. Mag. Tokyo 103: 325-334[PDF]

Kawai, H. and Nabata, S. 1990. Life history and systematic position of Pseudochorda gracilis sp. nov. (Laminariales, Phaeophyceae). J. Phycology 26: 721-727.

Kawai, H., Müller, D.G., Fölster, E. and Häder, D.-P. 1990. Phototactic responses in the gametes of a brown alga, Ectocarpus siliculosus. Planta 182: 292-297.h[PDF]

Müller, D.G., Kawai, H., Stache, B., Fölster, E. and Boland, B. 1990. Sexual pheromones and gamete chemotaxis in Analipus japonicus (Phaeophyceae). Experientia 46: 534-536.

Mimuro, M., Katoh, T. and Kawai, H. 1990. Spatial arrangement of pigments and their interaction in the fucoxanthin-chlorophyll a/c protein assembly (FCPA) isolated from the brown alga Dictyota dichotoma: Analysis by means of polarized spectroscopy. Biochimica et Biophysica Acta 1015: 450-456.[PDF]

Müller, D.G., Kawai H., Stache, B. and Lanka, S. 1990. A virus infection in the marine brown alga Ectocarpus siliculosus (Phaeophyceae). Botanica Acta 103(1): 72-82.


Kawai, H. 1989. First report of Phaeosaccion collinsii Farlow (Chrysophyceae, Sarcinochrysidales) from Japan. Jpn. J. Phycol. 37(3): 239-243.

Kawai, H. 1989. Life history and systematic position of Heteroralfsia saxicola gen. et comb. nov. (Ralfsiaceae, Phaeophyceae). Phycologia 28(2): 243-251.[PDF]

Kawai, H. and Inouye, I. 1989. Flagellar autofluorescence in forty-four chlorophyll c-containing algae. Phycologia 28(2): 222-227.[PDF]

Kawai, H. 1988. A flavin-like autofluorescent substance in the posterior flagellum of golden and brown algae. J. Phycol. 24(1): 114-117.

Yoshida, T. and Kawai, H. 1987. Taxonomic study of the genus Myagropsis (Cystoseiraceae, Phaeophyta). Bot Mag. Tokyo 100(2): 165-173.

Kawai, H. 1986. Life history and systematic position of Akkesiphycus lubricus (Phaeophyceae). J. Phycol. 22(4): 286-291.

Lim, B.-L., Kawai, H., Hori, H. and Osawa, S. 1986. Molecular evolution of 5S ribosomal RNA from red and brown algae. Jpn. J. Genet. 61(2):169-176.[PDF]

Kawai, H. and Kurogi, M. 1985. On the life history of Pseudochorda nagaii (Pseudochordaceae fam. nov.) and its transfer from Chordariales to Laminariales (Phaeophyta). Phycologia 24(3): 289-296.[PDF]

Kawai, H. 1983. Morphological observation on Acrothrix gracilis newly found in Japan. Jap. J. Phycol. 31(3): 167-172.

Kawai H. and Kurogi, M. 1983. Morphology and life history in culture of a new variety of Litosiphon groenlandicus from Japan. J. Fac. Sci., Hokkaido Univ. ser. V (Botany). 13(1): 25-38.[PDF]

Kawai H. and Kurogi, M. 1980. Morphological observation on a brown alga Delamarea attenuata, new to Japan. Jap. J. Phycol. 28(4): 225-231.


Kawai, H., Henry, E.C. 2016. ‘Phaeophyceae (Phaeophyta/Heterokonta/Ochrophyta)’ in ‘Handbook of the Protists’ Springer. In press

Kawai, H., Hanyuda, T., Uwai, S. 2016. ‘Evolution and biogeography of laminarialean kelps’ (Ed. Z. Hu and C. Fraser) in ‘Seaweed Phylogeography’. Springer. Pp. 227–249.

Nakayama, T., Kawai, H. 2015. ‘Introduction (Heterokontobionta p.p.), Cryptophyta, Dinophyta, Haptophyta, Heterkontophyta (except diatoms, Phaeophyceae, Eustigmatophyceae), Chlorarachniophyta, Euglenophyta’ (Ed. W. Frey) in ‘Syllabus of Plant Families – A. Engler’s Syllabus der Pflanzenfamilien Part 2/1 Photoautotrophic eukaryotic Algae.’ Borntraeger Verlagsbuchhandlung, Stuttgart.

Ni-Ni-Win, Hanyuda, T., Draisma, S.G.A., Lim, P.-E., Phang, S.-M., Kawai, H. 2013. Taxonomy of the genus Padina (Dictyotales, Phaeophyceae) based on morphological and molecular evidences, with key to species identification. In Taxonomy of Southeastern Asian Seaweeds II. Eds. Phang & Lim. pp. 119-174.

J. Mark Cock, Lieven Sterck, Sophia Ahmed, Andrew E. Allen, Grigoris Amoutzias, Veronique Anthouard, François Artiguenave, Alok Arun, Jean-Marc Aury, Jonathan H. Badger, Bank Beszteri, Kenny Billiau, Eric Bonnet, John H. Bothwell, Chris Bowler, Catherine Boyen, Colin Brownlee, Carl J. Carrano, Bénédicte Charrier, Ga Youn Cho, Susana M. Coelho, Jonas Collén, Gildas Le Corguillé, Erwan Corre, Laurence Dartevelle, Corinne Da Silva, Ludovic Delage, Nicolas Delaroque, Simon M. Dittami, Sylvie Doulbeau, Marek Elias, Garry Farnham, Claire M.M. Gachon, Olivier Godfroy, Bernhard Gschloessl, Svenja Heesch, Kamel Jabbari, Claire Jubin, Hiroshi Kawai, Kei Kimura, Bernard Kloareg, Frithjof C. Küpper, Daniel Lang, Aude Le Bail, Rémy Luthringer, Catherine Leblanc, Patrice Lerouge, Martin Lohr, Pascal J. Lopez, Nicolas Macaisne, Cindy Martens, Florian Maumus, Gurvan Michel, Diego Miranda-Saavedra, Julia Morales, Hervé Moreau, Taizo Motomura, Chikako Nagasato, Carolyn A. Napoli, David R. Nelson, Pi Nyvall-Collén, Akira F. Peters, Cyril Pommier, Philippe Potin, Julie Poulain, Hadi Quesneville, Betsy Read, Stefan A. Rensing, Andrés Ritter, Sylvie Rousvoal , Manoj Samanta, Gaelle Samson, Declan C. Schroeder, Delphine Scornet, Béatrice Ségurens, Martina Strittmatter, Thierry Tonon, James W. Tregear, Klaus Valentin, Peter von Dassow, Takahiro Yamagishi, Pierre Rouzé, Yves Van de Peer, Patrick Wincker (The Ectocarpus Genome Consortium). 2012. The Ectocarpus genome and brown algal genomics. In G. Piganeau (Ed.) Genomic Insights into the Biology of Algae. Advances in Botanical Research 64: 141-184 Elsevier Ltd.: Acadmic Press. ISBN: 9780123914996.

Lim P.E., Kawai, H., Phang, S.M. 2008. Some Ralfsiales from Malaysia. Pp. 77-81. In: ‘Taxonomy of Southeast Asian Seaweeds’. (eds. Phang, S.M., Lewmanomont, K., Lim P.E.) Univ. Malaya, Kuala Lumpur, Malaysia. 197. Pp.

Kawai, H., Henry, E.C. 2007. “Macroalgae” Pp. 33-36. In Sampling Biodiversity in Coastal Communities. NaGISA Protocols for Seagrass and Macroalgal Habitats. Eds Rigby, P.R., Iken, K., Shirayama, Y. Kyoto University Press. 145 pp.

Kawai, H., Motomura, T. and Okuda, K. 2005. Isolation and Purification Techniques for Macroalgae. In: Algal Culturing Techniques (ed. by R. A. Andersen), Elsevier Academic Press, Burlington, MA, U.S.A., pp. 133-143.

Shirayama, Y., Rowden, A. A. Gordon, D. P. Kawai, H. 2002. Latitudinal Biodiversity in coastal macrophyte communities. pp. 164-182. In Biodiversity Research Methods: IBOY in Western Pacific and Asia. Eds. Nakashizuka, T. and Stork, N. Kyoto University Press. 216 pp.

Kawai, H., Kreimer, G. 2000. Sensory mechanisms: Phototaxis and light perception in algae. pp. 124-146. In The Flagellates: Unity, diversity and evolution. Eds. B.S.C. Leadbeater. & J. Green. Taylor & Francis, London. 401 pp.